1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega‐Science Institute of Oceanology, Chinese Academy of Sciences Qingdao China
2. Laboratory for Marine Biology and Biotechnology Qingdao National Laboratory for Marine Science and Technology Qingdao China
3. Liaoning Ocean and Fisheries Science Research Institute, Liaoning Province Key Laboratory of Marine Biological Resources and Ecology Dalian Key Laboratory of Conservation of Fishery Resources Dalian China