1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650204 China
2. Institute of Alpine Economic Plants; Yunnan Academy of Agricultural Sciences; Lijiang 674100 Yunnan China
3. Faculty of Biology; Vietnam National University − University of Science; Hanoi Vietnam
4. Germplasm Bank of Wild Species in Southwest China; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650204 China
5. University of the Chinese Academy of Sciences; Beijing 100049 China
6. Royal Botanic Garden Edinburgh; Edinburgh EH3 5LR UK
7. Vietnam National Museum of Nature; Vietnam Academy of Science and Technology; Hanoi Vietnam