1. Center for Macroecology; Evolution and Climate; Natural History Museum of Denmark; University of Copenhagen; Copenhagen Ø DK-2100 Denmark
2. Department of Biodiversity Informatics and Genetics; Swedish Museum of Natural History; Stockholm 10405 Sweden
3. Department of Vertebrate Zoology; Swedish Museum of Natural History; Stockholm 10405 Sweden
4. Center for Macroecology; Evolution and Climate; University of Copenhagen; Copenhagen Ø DK-2100 Denmark
5. Quebec Centre for Biodiversity Science; Department of Biology; McGill University; Montreal QC H3A-1B1 Canada
6. Department of Life Sciences; Imperial College London; Silwood Park campus Ascot SL5 7PY UK