1. Environmental Biology Department Institute of Environmental Sciences CML, Leiden University Einsteinweg 2 2333 CC Leiden the Netherlands
2. South China Botanical Garden Chinese Academy of Sciences No.723 Xingke Road, Tianhe District510650 Guangzhou China
3. Institute of Ecology and Earth Sciences University of Tartu Ülikooli 18 50090 Tartu Estonia
4. School of Biological Sciences Faculty of Science University of Western Australia 35 Stirling Highway Perth WA 6009 Australia
5. Understanding Evolution Group Naturalis Biodiversity Center 2332 AA Leiden the Netherlands
6. Department of Evolutionary and Population Biology Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam 1098 XHAmsterdam the Netherlands