1. Department of Anatomy; University of Otago; PO Box 913 Dunedin 9054 New Zealand
2. Department of Pathology; University of Otago; Christchurch 8140 New Zealand
3. Department of Public Health and General Practice; University of Otago; Christchurch 8140 New Zealand
4. Génétique Quantitative et Méthologie de la Sélection; UMR de Génétique Végétale; INRA - Université Paris-Sud - CNRS - AgroParisTech; Ferme du Moulon; 91190 Gif-sur-Yvette France
5. Department of Conservation; Marine Species and Threats Team; Wellington 6011 New Zealand