1. College of Fisheries and Life Science Dalian Ocean University Dalian China
2. Key Laboratory of South China Sea Fishery Resources Exploitation and Utilization Ministry of Agriculture and Rural Affairs, South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences Guangzhou China
3. Sanya Tropical Fisheries Research Institute Sanya China
4. Key Laboratory of Aquatic Genomics, Ministry of Agriculture and Rural Affairs Chinese Academy of Fishery Science Beijing China