1. Evolutionary Ecology and Genetics group; Biodiversity Research Centre; Earth and Life Institute; Académie Louvain; Croix du Sud 4 1348 Louvain-la-Neuve Belgium
2. Université Montpellier 2; CNRS; Institut des Sciences de l'Evolution; CC65, Place Eugène Bataillon 34095 Montpellier France
3. Terrestrial Ecology Unit; Ghent University; Ghent Belgium