1. College of Veterinary Medicine Yangzhou University Yangzhou China
2. Jiangsu Co‐Innovation Center for the Prevention and Control of Important Animal Infectious Disease and Zoonoses Yangzhou China
3. Jiangsu Research Centre of Engineering and Technology for Prevention and Control of Poultry Disease Yangzhou China
4. Department of Avian Infectious Diseases Shanghai Veterinary Research Institute Chinese Academy of Agricultural Science Shanghai China
5. The International Joint Laboratory for Cooperation in Agriculture and Agricultural Product Safety Ministry of Education Yangzhou University Yangzhou China