1. AIMS@JCU, Australian Institute of Marine Science & College of Marine and Environmental Sciences; James Cook University; Townsville Qld 4811 Australia
2. Centre for Sustainable Tropical Fisheries and Aquaculture & College of Marine and Environmental Sciences; James Cook University; Townsville Qld 4811 Australia
3. Australian Institute of Marine Science; Townsville Qld 4810 Australia
4. Great Lakes Institute for Environmental Research, Department of Earth and Environmental Sciences; University of Windsor; Windsor ON N9B 3P4 Canada