1. Miyako Laboratory; Tohoku National Fisheries Research Institute; Japan Fisheries Research and Education Agency; 4-9-1, Sakiyama Miyako Iwate 027-0097 Japan
2. Ishinomaki Sensyu University; 1, Shinmito, Minamisakai Ishinomaki Miyagi 986-8580 Japan
3. Tohoku National Fisheries Research Institute; Japan Fisheries Research and Education Agency; 3-27-5, Shinhama Shiogama Miyagi 985-0001 Japan
4. International Coastal Research Center; Atmosphere and Ocean Research Institute; The University of Tokyo; 2-106-1, Akahama Otsuchi Iwate 028-1102 Japan