1. Sugadaira Montane Research Center; University of Tsukuba; Nagano; Japan
2. Entomology Group, Institut für Spezielle Zoologie und Evolutionsbiologie mit Phyletischem Museum; FSU Jena; Germany
3. Laboratory of Systematic Entomology, Department of Ecology and Systematics; Graduate School of Agriculture, Hokkaido University; Sapporo; Japan