1. National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, College of Horticulture & Forestry Sciences Huazhong Agricultural University Wuhan 430070 China
2. Hubei Hongshan Laboratory Wuhan 430070 China
3. Department of Biosciences Durham University South Road Durham DH1 3LE UK
4. Endomembrane Structure and Function Research Group, Department of Biological and Medical Sciences Oxford Brookes University Oxford OX3 0BP UK