1. Department of Cell and Systems Biology; University of Toronto; 25 Willcocks Street Toronto ON M5S 3B2 Canada
2. Agriculture and Agri-Food Canada/Agriculture et Agroalimentaire Canada; KW Neatby bldg; 960 Carling Ave. Ottawa ON K1A 0C6 Canada
3. Centre for the Analysis of Genome Function and Evolution; University of Toronto; 25 Willcocks Street Toronto ON M5S 3B2 Canada