1. Aix Marseille Univ; Univ Avignon; CNRS, IRD, IMBE, Case 421 Av Escadrille Normandie Niémen 13 397 Marseille cedex 20 France
2. Department of Biology; McGill University; Stewart Biology Building 1205 Docteur Penfield Montreal QC H3A 1B1 Canada
3. Département des sciences naturelles, Université du Québec en Outaouais; Institut des sciences de la forêt tempérée; 58 Rue Principale Ripon QC J0V 1V0 Canada
4. Apex Resource Management Solutions Ltd.; 937 Kingsmere Ave Ottawa, ON K2A 3K2 Canada
5. Quebec Centre for Biodiversity Science; Stewart Biology Building 1205 Docteur Penfield Montreal QC H3A 1B1 Canada