1. School of Biological Sciences, Gehrmann Building; The University of Queensland and Australian Research Council Centre of Excellence for Coral Reef Studies; St Lucia Queensland 4072 Australia
2. Centre for Integrative Ecology, School of Life and Environmental Sciences; Deakin University; Burwood Victoria 3125 Australia
3. Centre for Sustainable Tropical Fisheries and Aquaculture, School of Earth and Environmental Sciences; James Cook University; Townsville Queensland 4810 Australia