1. Department of Biology, Graduate Program in Ecology and Evolutionary Biology University of Oklahoma 730 Van Vleet Oval Norman OK 73019 U.S.A.
2. Department of Biology, Graduate Program in Ecology and Evolutionary Biology University of Louisville 2301 S 3rd St, Louisville KY 40208 U.S.A.
3. Center for Conservation and Sustainability, Smithsonian Conservation Biology Institute National Zoological Park P.O. Box 37012, MRC 705 Washington D.C. 20013‐7012 U.S.A.