1. Marwell Wildlife, Colden Common Winchester UK
2. Faculty of Science of Tunis, Research Laboratory of Biodiversity, Management and Conservation of Biological Systems University of Tunis El Manar Tunis Tunisia
3. Direction Générale des Forêts Ministre de l'Agriculture, des Ressources Hydrauliques et de la Pêche Maritime Tunis Tunisia
4. School of Biological Sciences, Faculty of Environmental and Life Sciences University of Southampton Southampton UK