1. State Key Laboratory of Developmental Biology of Freshwater Fish; Hunan Normal University; Changsha Hunan 410081 China
2. College of Life Sciences; Hunan Normal University; Changsha Hunan 410081 China
3. Key Laboratory of Freshwater Aquatic Genetic Resources, Ministry of Agriculture; Shanghai Ocean University; Shanghai 201306 China
4. National Demonstration Center for Experimental Fisheries Science Education; Shanghai Ocean University; Shanghai 201306 China
5. Shanghai Engineering Research Center of Aquaculture; Shanghai Ocean University; Shanghai 201306 China