1. Key Laboratory of Freshwater Aquatic Genetic Resources, Ministry of Agriculture; Shanghai Ocean University; Shanghai 201306 China
2. National Demonstration Center for Experimental Fisheries Science Education; Shanghai Ocean University; Shanghai 201306 China
3. Shanghai Engineering Research Center of Aquaculture; Shanghai Ocean University; Shanghai 201306 China
4. Key Laboratory of Freshwater Fisheries and Germplasm Resources Utilization, Ministry of Agriculture, Freshwater Fisheries Research Center, Chinese Academy of Fishery Sciences; Wuxi 214081 China
5. Freshwater Fisheries Research Institute of Jiangsu Province; Nanjing 210017 China
6. Panjin Guanghe Crab Industry Co. Ltd; Panjin China