1. Department of Bioscience and Biotechnology Fukui Prefectural University 4‐1‐1 Matsuoka‐Kenjojima Eiheiji Fukui 910‐1195 Japan
2. Plant Nutrition and Crop Physiology Department of Crop Science University of Göttingen Carl‐Sprengel‐Weg 1 Göttingen 37075 Germany
3. Institute of Plant Science and Resources Okayama University 2‐20‐1 Chuo Kurashiki Okayama 710‐0046 Japan
4. Department of Ecophysiology Institute of Cellular Botany University of Bonn Kirschallee 1 53115 Germany
5. Faculty of Agro‐Food Science Niigata Agro‐Food University 2416 Hiranedai Tainai Niigata 959‐2702 Japan