1. Faculty of Fisheries Sciences Hokkaido University Hakodate Japan
2. Doto Research Branch Salmon and Freshwater Fisheries Research Institute, Hokkaido Research Organization Nakashibetsu Japan
3. Salmon and Freshwater Fisheries Research Institute, Hokkaido Research Organization Eniwa Japan
4. Kitami Salmon Enhancement Program Association Abashiri Japan
5. Abashiri Fisheries Cooperative Abashiri Japan