1. Institute of Ecology and Earth Sciences; University of Tartu; Lai 40 Tartu 51005 Estonia
2. Evolution & Ecology Research Centre; School of Biological, Earth and Environmental Sciences; University of New South Wales; Sydney NSW 2052 Australia
3. Pyrenean Institute of Ecology; Spanish National Research Council (IPE-CSIC); Av. Montañana 1005 Zaragoza 50059 Spain
4. School of Plant Biology; University of Western Australia; Crawley Perth WA 6009 Australia
5. Institute for Land; Water and Society; School of Environmental Science; Charles Sturt University; Albury NSW 2640 Australia