1. Finnish Centre of Excellence in Metapopulation Biology Department of Biosciences P.O. Box 65, FI‐00014 University of Helsinki Finland
2. School of Life Sciences University of KwaZulu‐Natal Durban 4041 South Africa
3. Department of Geography University of Cambridge Downing Place Cambridge CB2 3EN United Kingdom
4. Ezemvelo KwaZulu‐Natal Wildlife P.O. Box 13053, Cascades 3202 South Africa