1. School of Biological Sciences; The University of Queensland; Brisbane QLD 4072 Australia
2. School of Geography; Planning, and Environmental Management; The University of Queensland; Brisbane QLD 4072 Australia
3. CSIRO Land and Water; 41 Boggo Road Dutton Park QLD 4102 Australia
4. School of Mathematics and Physics; The University of Queensland; Brisbane QLD 4072 Australia
5. Department of Life Sciences; Imperial College London; Silwood Park, Ascot SL5 7PY Berkshire England United Kingdom