1. IRD; UMR DIADE; BP 64501 34394 Montpellier Cedex 5 France
2. UM2 and IRD; UMR AMAP; Boulevard de la Lironde, PS 2 34398 Montpellier France
3. IRD; UMR AMAP; Herbarium NOU 98848 Nouméa New Caledonia
4. IRD; UMR DIADE; 98848 Nouméa New Caledonia
5. Tropical Conservation Biology and Environmental Science; University of Hawai'i at Hilo; 200 West Kawili St. Hilo HI 96720 USA