1. Department of Genetics, Evolution and Environment, Center for Biodiversity and Environment Research; University College of London; London U.K.
2. Leibniz-Institute of Freshwater Ecology and Inland Fisheries (IGB); Müggelseedamm 310 12587 Berlin Germany
3. Department of Biology, Chemistry, Pharmacy, Institute of Biology; Freie Universität Berlin; Königin-Luise-Strasse 1-3 14195 Berlin Germany