1. Center for Systems Integration and Sustainability, Department of Fisheries and Wildlife; Michigan State University; East Lansing MI 48823 U.S.A.
2. Department of Ecology and Evolutionary Biology; Yale University; New Haven CT 06520 U.S.A.
3. Conservation International; Arlington VA 22202 U.S.A.
4. Department of Geography; The University of North Carolina at Chapel Hill; Chapel Hill NC 27599 U.S.A.
5. Department of Geography; Michigan State University; East Lansing MI 48824 U.S.A.