1. Institute of Ecology and Botany Centre for Ecological Research Vácrátót H‐2163 Hungary
2. Estación Biológica de Doñana‐CSIC Seville 41092 Spain
3. MTA‐ELTE, Theoretical Biology and Evolutionary Ecology Research Group, Department of Plant Systematics, Ecology and Theoretical Biology Eötvös Loránd University Budapest H‐1053 Hungary
4. Centre for Evolutionary Biology, School of Biological Sciences University of Western Australia Crawley WA 6009 Australia