1. Department of Botany; Institute of Biological Sciences Universidade Federal de Minas Gerais; Belo Horizonte Brazil
2. Institute of Biological and Health Sciences, Universidade Federal Rural do Rio de Janeiro; Seropédica Brazil
3. Institute of Forest Engineering, Universidade Federal Rural do Rio de Janeiro; Seropédica Brazil
4. Department of Botany; Institute of Biological and Health Sciences, Universidade Federal Rural do Rio de Janeiro; Seropédica Brazil
5. Rio de Janeiro Botanical Garden; Rio de Janeiro Brazil