1. School of Plant Biology and Institute of Agriculture; The University of Western Australia; 35 Stirling Highway Perth Western Australia 6009 Australia
2. Future Farm Industries Cooperative Research Centre; The University of Western Australia; 35 Stirling Highway Perth Western Australia 6009 Australia
3. School of Earth and Environment; The University of Western Australia; 35 Stirling Highway Perth Western Australia 6009 Australia
4. Cranfield Soil and AgriFood Institute; Department of Environmental Science and Technology; School of Applied Sciences; Cranfield University; Cranfield MK43 0SZ UK