1. State Key Laboratory of Vegetation and Environmental Change Institute of Botany, Chinese Academy of Sciences Xiangshan Beijing 100093 China
2. University of Chinese Academy of Sciences Yuquan Road Beijing 100049 China
3. School of Environmental Science and Engineering Nanjing University of Information Science and Technology Nanjing 210044 China
4. Institute of Agricultural Resources and Regional Planning Chinese Academy of Agricultural Sciences Beijing 100093 China
5. School of Biological Sciences The University of Hong Kong Hong Kong 999077 China