1. Institute of Digestive Disease Chinese University of Hong Kong Hong Kong China
2. Division of Gastroenterology and Hepatology, Department of Medicine and Therapeutics Prince of Wales Hospital Hong Kong China
3. Translational Gastroenterology Unit, John Radcliffe hospital University of Oxford Oxford UK
4. Wellcome Centre for Human Genetics, Nuffield Department of Medicine University of Oxford Oxford UK
5. Department of Clinical Oncology Chinese University of Hong Kong Hong Kong China