1. Division of Mucosal Immunology; Department of Microbiology and Immunology; Institute of Medical Science; The University of Tokyo; Tokyo Japan
2. Core Research for Evolutional Science and Technology (CREST); Japan Science and Technology Agency; Tokyo Japan
3. Department of Medical Genome Sciences, Graduate School of Frontier Sciences; The University of Tokyo; Chiba Japan
4. Laboratory of Vaccine Materials; National Institute of Biomedical Innovation; Osaka Japan
5. International Research and Development Center for Mucosal Vaccines, Institute of Medical Science; The University of Tokyo; Tokyo Japan
6. Graduate School of Pharmaceutical Sciences; Osaka University; Osaka Japan