1. Department of Biology; The Pennsylvania State University; University Park PA 16802 USA
2. Papahānaumokuākea Marine National Monument; Office of the National Marine Sanctuaries; National Oceanic and Atmospheric Administration; Honolulu HI 96825 USA
3. Hawai'i Institute of Marine Biology; School of Ocean and Earth Science and Technology; University of Hawai'i at Mānoa; Kāne'ohe HI 96744 USA
4. Department of Biology; University of Hawai'i at Mānoa; Honolulu HI 96822 USA