1. School of Geographical and Earth Sciences; University of Glasgow; Glasgow G12 8QQ UK
2. Department of Earth Sciences; University of St Andrews; Fife KY16 9AJ UK
3. Marine Biology and Ecology Research Centre; School of Marine Science and Engineering; Plymouth University; Drake Circus Plymouth PL4 8AA UK
4. Plymouth Marine Laboratory; Prospect Place Plymouth PL1 3DH UK
5. CNRS, Laboratoire Adaptation et Diverstié en Milieu Marin, Station Biologique de Roscoff; Place Georges Teissier 29688 Roscoff Cedex France
6. Université Pierre et Marie Curie - Paris 6; Laboratoire Adaptation et Diversité en Milieu Marin; Station Biologique de Roscoff; Place Georges Teissier 29688 Roscoff Cedex France