1. The United Graduate School of Agricultural Sciences; Kagoshima University; Kagoshima Japan
2. National Agriculture and Food Research Organization (NARO); Institute of Livestock and Grassland Science; Nasushiobara Tochigi Japan
3. Japan International Research Center for Agricultural Sciences (JIRCAS); Tsukuba Ibaraki Japan
4. Faculty of Agriculture; University of the Ryukyus; Nishihara Okinawa Japan