1. Sustainable Agri-Food Sciences Division; Agri-Food & Biosciences Institute; Loughgall Co. Armagh BT61 8JA UK
2. School of Biological Sciences; Queen's University Belfast; Belfast BT9 7PL UK
3. Sustainable Agri-Food Sciences Division; Agri-Food & Biosciences Institute; Newforge Lane Belfast BT9 5PX UK
4. Department of Plant, Soil and Microbial Science; Michigan State University; East Lansing MI 48824 USA
5. ARS/PSI-Genetic Improvement of Fruits and Vegetables Laboratory; USDA; Beltsville MD 20705 USA