1. School of Plant Sciences; Haramaya University; Dire Dawa Ethiopia
2. Department of Zoology and Entomology; University of Fort Hare; Alice South Africa
3. Department of Crop Production; University of Swaziland; Luyengo Campus Swaziland
4. Department of Biological and Environmental Sciences; Walter Sisulu University; Mthatha South Africa
5. IRD/CRNS UMR IRD 247 EGCE; Laboratoire Evolution Génomes Comportement et Ecologie; Comportement et Ecologie; CNRS; Gif sur Yvette Cedex France
6. Université Paris - Sud 11; Orsay Cedex France