1. Department of Botany, Faculty of Science, Kasetsart University Bangkok Thailand
2. Hala‐Bala Wilflife Research Station (Narathiwat), Department of National Park, Wildlife and Plant conservation Bangkok Thailand
3. Kasetsart Agricultural and Agro‐Industrial Product Improvement Institute, Kasetsart University Bangkok Thailand
4. Botany Department, School of Natural Sciences, Trinity College Dublin Dublin Ireland