1. Arizona State University Tempe AZ USA
2. University of Nevada Reno NV USA
3. Smithsonian Tropical Research Institute Panama City Panama
4. Queen Mary University London London UK
5. University of Queensland Brisbane QLD Australia
6. Georgia Southern University Statesboro GA USA
7. University of California, Los Angeles Los Angeles CA USA
8. Harvard University Cambridge MA USA
9. Northeastern University Boston MA USA
10. University of Michigan Ann Arbor MI USA
11. Washington University Saint Louis MO USA
12. Florida International University Miami FL USA