1. Does bird predation influence the spot-number variation in Maniola jurtina (Lepidoptera);Bengtson;Biological Journal of the Linnean Society,1981
2. Brakefield , P. M. 1979 An experimental study of the maintenance of variation in spot pattern in Maniola jurtina. Ph.D. Thesis University of Liverpool
3. The ecological genetics of quantitative characters of Maniola jurtina and other butterflies;Brakefield;The Biology of Butterflies,1984