1. UMR CNRS 5554 and UMR IRD 226; Institut des Sciences de l'Evolution de Montpellier; Université Montpellier II; Place Eugène Bataillon, CC065; 34095; Montpellier, Cedex 05; France
2. UMR CNRS-MNHN 7205; Origine, Structure, Evolution de la Biodiversité; 55 rue Buffon; 75005; Paris; France
3. Ural Division of the Russian Academy of Sciences; Institute of Plant and Animal Ecology; ul. 8 Marta; 202; Ekaterinburg; 620144; Russia
4. UMR CNRS 6282 Biogéosciences; Université de Bourgogne; 6 Boulevard Gabriel; 21000; Dijon; France
5. Severtsov Institute of Ecology and Evolution; Russian Academy of Sciences; Leninskii pr 33; 117071; Moscow; Russia
6. INRA, UMR CBGP (Inra/Ird/Cirad/Montpellier SupAgro); Campus international de Baillarguet, CS 30016; 34988; Montferrier-sur-Lez cedex; France