1. Department of Entomology; North Carolina State University; Raleigh NC 27695 USA
2. Smithsonian Tropical Research Institute; Apartado 0843-03092 Ancón República de Panamá
3. Departamento de Ciencias Naturales; Universidad Técnica Particular de Loja; Loja CP 11-01-608 Ecuador
4. Graduate Program in Ecology and Evolutionary Biology and Department of Zoology; University of Oklahoma; Norman OK 73019 USA