1. Department of Animal and Plant Sciences; University of Sheffield, Western Bank; Sheffield S10 2TN UK
2. Botany Department; Rhodes University; Grahamstown 6139 South Africa
3. CNRS, Université de Toulouse, ENFA; UMR5174 EDB (Laboratoire Évolution and Diversité Biologique); 118 Route de Narbonne Toulouse 31062 France
4. Department of Primary Industry and Fisheries; Northern Territory Government; Darwin NT 0801 Australia
5. School of Chemistry and Biochemistry (honorary research fellow); University of Western Australia; Crawley WA 6009 Australia
6. Beacon Bay; East London 5205 South Africa