1. Department of Biological Sciences, Graduate School of ScienceOsaka University Machikaneyama‐cho 1‐1, Toyonaka Osaka 560‐0043 Japan
2. Agricultural Genetics InstituteNational Key Laboratory for Plant Cell Biotechnology Pham Van Dong road, Bac Tu Liem district Ha Noi Vietnam
3. Institute of Plant Science and ResourcesOkayama University Kurashiki Okayama 710‐0046 Japan