1. Institute of Biodiversity, Animal Health, and Comparative Medicine; University of Glasgow; Graham Kerr Building Glasgow G12 8QQ UK
2. Department of Zoology; University of Cambridge; Downing Street Cambridge CB2 3EJ UK
3. Centre d'Ecologie Fonctionnelle et Evolutive CEFE UMR 5175; CNRS - Université de Montpellier - Université Paul-Valéry Montpellier - EPHE; 919 route de Mende 34293 Montpellier Cedex 5 France
4. MARE - Marine and Environmental Sciences Centre; ISPA - Instituto Universitário; Rua Jardim do Tabaco 34 1149-041 Lisbon Portugal