1. Laboratory of Aquatic Ecology, Evolution and Conservation; University of Leuven; Deberiotstraat 32 3000 Leuven Belgium
2. Research Institute for Nature and Forest; Gaverstraat 4 B-9500 Geraardsbergen Belgium
3. Laboratory of Socioecology and Social Evolution; Department of Biology; University of Leuven; Naamsestraat 59 3000 Leuven Belgium
4. Laboratory of Forensic Genetics and Molecular Archaeology; University of Leuven; Kapucijnenvoer 33 3000 Leuven Belgium
5. Station d'Ecologie Expérimentale du CNRS à Moulis; USR 2936 09200 Moulis France
6. Department of Genetics; University of Leicester; University Road LE1 7RH Leicester UK