1. Department of Biology; Center for Conservation Biology; Stanford University; Stanford CA 94305 USA
2. Copal de Agua Buena; Coto Brus 8257 Costa Rica
3. The Natural Capital Project; Stanford CA 94305 USA
4. Department of Biology; Stanford University; Stanford CA 94305 USA
5. Department of Biology; Center for Conservation Biology; Woods Institute for the Environment; Stanford University; Stanford CA 94305 USA