1. Key Laboratory of Tropical Marine Bio‐resources and Ecology (LMB), Guangdong Provincial Key Laboratory of Applied Marine Biology (LAMB) South China Sea Institute of Oceanology, Chinese Academy of Sciences Guangzhou China
2. University of Chinese Academy of Sciences Beijing China
3. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou China
4. Department of Marine Biology Texas A&M University at Galveston Galveston Texas USA
5. Key Laboratory of Science and Technology on Operational Oceanography South China Sea Institute of Oceanology, Chinese Academy of Sciences Guangzhou China