1. Dipartimento di Scienze; Università degli studi Roma Tre; Viale Guglielmo Marconi 446; 00146 Roma Italy
2. Department of Applied and Environmental Biology; Rivers State University of Science and Technology, PMB 5080; Port Harcourt Rivers State Nigeria
3. Institute for Development, Ecology, Conservation and Cooperation; via G. Tomasi di Lampedusa 33; I-00144 Rome Italy